Despite plenty of data in insects shape variation, papers on environmental factors effect on such variation are scarce and in ground beetles are practically absent. The aim of this research was to: (i) model the effect of the region of habitation, urbanization and habitat vegetation into the shape variation in widespread carabid species; (ii) describe sexual shape dimorphism in studied species. Samples were pitfall trapped in different regions of its area in the spectrum of anthropogenic influence (cities, suburbs, natural biotopes). One thousand and one hundred sixty-eight specimens were analyzed for six morphometric traits and terminal points of those measurements were used as landmarks for Procrustes analysis. We used linear models to reveal which factor (region, urbanization or vegetation) and in what direction affected beetles shape. Results showed that males in C. granulatus had more convex elytra and head, and the more convex in apical-basal direction pronotum. In its area from the east to the west elytra flattened in medial-distal direction and pronotum flattened in apical-distal direction. In disturbed urban environment beetles pronotum became more convex in both sexes and males elytra became more convex too. In open habitats (meadows, lawns) beetles became more flattened.

Despite plenty of data in insects shape variation, papers on environmental factors effect on such variation are scarce and in ground beetles are practically absent. The aim of this research was to: (i) model the effect of the region of habitation, urbanization and habitat vegetation into the shape variation in widespread carabid species; (ii) describe sexual shape dimorphism in studied species. Samples were pitfall trapped in different regions of its area in the spectrum of anthropogenic influence (cities, suburbs, natural biotopes). One thousand and one hundred sixty-eight specimens were analyzed for six morphometric traits and terminal points of those measurements were used as landmarks for Procrustes analysis. We used linear models to reveal which factor (region, urbanization or vegetation) and in what direction affected beetles shape. Results showed that males in C. granulatus had more convex elytra and head, and the more convex in apical-basal direction pronotum. In its area from the east to the west elytra flattened in medial-distal direction and pronotum flattened in apical-distal direction. In disturbed urban environment beetles pronotum became more convex in both sexes and males elytra became more convex too. In open habitats (meadows, lawns) beetles became more flattened.