1CAS Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China 2CAS Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China 3State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China 4State Key Laboratory of Mycology (SKLM), Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China 5CAS Key Laboratory of Aquatic Biodiversity and Conservation, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China 6Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, China 7Department of Marine Organism Taxonomy & Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China 8CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China 9CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China 10State Key Laboratory of Genetic Resources and Evolution & Yunnan Key Laboratory of Biodiversity and Ecological Security of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650221, China 11CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China 12CAS Key Laboratory of Plant Resource Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China 13CAS Key Laboratory of Wetland Ecology and Environment, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China 14CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China 15State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China 16National Center for Mathematics and Interdisciplinary Sciences, Key Laboratory of Random Complex Structures and Data Science, Academy of Mathematics and Systems Science，Chinese Academy of Sciences, Beijing 100190, China 17Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China 18Institute of Entomology, College of Life Sciences, Nankai University, Tianjin 300071, China 19Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China 20Institute of Plant Protection, Beijing Academy of Agriculture and Forestry Sciences, Beijing 100097, China 21School of Ecology, Sun Yat-sen University, Shenzhen 518000, China 22School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China 23College of Plant Protection, Nanjing Agricultural University, Nanjing 210095, China 24The Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding 071002, China 25College of Life Sciences, Hunan Normal University, Changsha 410081, China 26College of Life Sciences, Capital Normal University, Beijing 100048, China 27School of Life Sciences, Nanjing Normal University, Nanjing 210046, China 28Department of Zoology, College of Life Sciences, Shaanxi Normal University, Xi’an 710119, China 29Institute of Insect Science and Technology, School of Life Sciences, South China Normal University, Guangzhou 510631, China 30College of Life Sciences, Shanghai Normal University, Shanghai 200234, China 31Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, College of Plant Protection, Northwest A&F University, Yangling 712100, China 32State Key Laboratory of Ecological Pest Control for Fujian and Taiwan Crops, College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou 350002, China 33Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, China 34Institute of Entomology, Guizhou University, Guiyang 550025, China 35School of Life Sciences, Southwest University, Chongqing 400715, China 36College of Life Science and Technology, Xinjiang University, Urumqi 830017, China 37College of Advanced Agricultural Sciences, University of Chinese Academy of Sciences, Beijing 100049, China 38College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China

1CAS Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China 2CAS Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China 3State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China 4State Key Laboratory of Mycology (SKLM), Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China 5CAS Key Laboratory of Aquatic Biodiversity and Conservation, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China 6Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, China 7Department of Marine Organism Taxonomy & Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China 8CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China 9CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China 10State Key Laboratory of Genetic Resources and Evolution & Yunnan Key Laboratory of Biodiversity and Ecological Security of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650221, China 11CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China 12CAS Key Laboratory of Plant Resource Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China 13CAS Key Laboratory of Wetland Ecology and Environment, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China 14CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, China 15State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China 16National Center for Mathematics and Interdisciplinary Sciences, Key Laboratory of Random Complex Structures and Data Science, Academy of Mathematics and Systems Science，Chinese Academy of Sciences, Beijing 100190, China 17Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China 18Institute of Entomology, College of Life Sciences, Nankai University, Tianjin 300071, China 19Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China 20Institute of Plant Protection, Beijing Academy of Agriculture and Forestry Sciences, Beijing 100097, China 21School of Ecology, Sun Yat-sen University, Shenzhen 518000, China 22School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China 23College of Plant Protection, Nanjing Agricultural University, Nanjing 210095, China 24The Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding 071002, China 25College of Life Sciences, Hunan Normal University, Changsha 410081, China 26College of Life Sciences, Capital Normal University, Beijing 100048, China 27School of Life Sciences, Nanjing Normal University, Nanjing 210046, China 28Department of Zoology, College of Life Sciences, Shaanxi Normal University, Xi’an 710119, China 29Institute of Insect Science and Technology, School of Life Sciences, South China Normal University, Guangzhou 510631, China 30College of Life Sciences, Shanghai Normal University, Shanghai 200234, China 31Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, College of Plant Protection, Northwest A&F University, Yangling 712100, China 32State Key Laboratory of Ecological Pest Control for Fujian and Taiwan Crops, College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou 350002, China 33Department of Entomology, College of Plant Protection, South China Agricultural University, Guangzhou 510642, China 34Institute of Entomology, Guizhou University, Guiyang 550025, China 35School of Life Sciences, Southwest University, Chongqing 400715, China 36College of Life Science and Technology, Xinjiang University, Urumqi 830017, China 37College of Advanced Agricultural Sciences, University of Chinese Academy of Sciences, Beijing 100049, China 38College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China